Dr. rer. nat. Ivan Calandra

Wissenschaftlicher Mitarbeiter


  • Controlled experiments & quantitative artifact microwear analysis
  • Dental evolution of terrestrial mammals through analyses of form and function
  • Keywords: archeology, climate, evolution, paleobiology, traceology


Professional appointments

  • Since 2021
    Head of IMPALA, Imaging Platform at LEIZA, Neuwied & Mainz, Germany.

  • 2016-2021
    Post-doctoral fellow, TraCEr lab, MONREPOS Archaeological Research Centre, RGZM, Neuwied, Germany.

  • 2016
    Scientific Mediator (science communication), University of Reims Champagne-Ardenne, Reims, France.

  • 2014-2015
    ATER (temporary teaching and research assistant), University of Reims Champagne-Ardenne, GEGENAA - EA 3795, Reims, France.

  • 2013-2014
    ATER, University of Burgundy Franche-Comté, UMR 6249 Chrono-Environnement, Besançon-Montbéliard, France.

  • 2013
    Study engineer, University of Burgundy Franche-Comté, UMR 6282 Biogéosciences, Dijon, France.

  • 2011-2012
    Post-doctoral fellow, University of Burgundy Franche-Comté, UMR 6282 Biogéosciences, Dijon, France.


  • 2008-2011
    PhD in Zoology. Zoological Institute and Museum (now Center of Natural History, CeNak), University Hamburg, Germany.

  • 2006-2008
    MSc in Paleontology, Sedimentology, and Paleoenvironments. University Claude Bernard Lyon 1 & École Normale Supérieure, Lyon, France.

  • 2003-06
    BSc in Geology. University Claude Bernard Lyon 1, France & University of Oregon, Eugene, USA.

Further Links


 Leibniz-Zentrum für Archäologie
Ludwig-Lindenschmit-Forum 1

56567 Neuwied
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  • Schunk L, Gneisinger W, Calandra I, Marreiros J. 2023. The role of artificial contact materials in experimental use-wear studies: A controlled proxy to understand use-wear polish formation. Journal of Archaeological Science: Reports 47:103737. DOI: https://doi.org/10.1016/j.jasrep.2022.103737.


  • Calandra I. 2022. A workflow for quality control in surface texture analysis applied to teeth and tools. Journal of Archaeological Science: Reports 46:103692. DOI: https://doi.org/10.1016/j.jasrep.2022.103692.
  • Calandra I, Bob K, Merceron G, Blateyron F, Hildebrandt A, Schulz-Kornas E, Souron A, Winkler DE. 2022. Surface texture analysis in Toothfrax and MountainsMap® SSFA module: Different software packages, different results? Peer Community Journal 2. DOI: https://doi.org/10.24072/pcjournal.204.


  • Malinsky-Buller A, Glauberman P, Ollivier V, Lauer T, Timms R, Frahm E, Brittingham A, Triller B, Kindler L, Knul MV, Krakovsky M, Joannin S, Hren MT, Bellier O, Clark AA, Blockley SPE, Arakelyan D, Marreiros J, Paixaco E, Calandra I, Ghukasyan R, Nora D, Nir N, Adigyozalyan A, Haydosyan H, Gasparyan B. 2021. Short-term occupations at high elevation during the Middle Paleolithic at Kalavan 2 (Republic of Armenia). PLOS ONE 16:e0245700. DOI: https://doi.org/10.1371/journal.pone.0245700.


  • Calandra I. 2020. Artefakte im Experiment. Archäologie in Deutschland 2020:20–21.
  • Calandra I, Gneisinger W, Marreiros J, Pedergnana A, Schunk L. 2020. Spurensuche im Mikrobereich. Archäologie in Deutschland 2020:22–25.
  • Calandra I, Gneisinger W, Marreiros J. 2020. A versatile mechanized setup for controlled experiments in archeology. STAR: Science & Technology of Archaeological Research 6:30–40. DOI: https://doi.org/10.1080/20548923.2020.1757899.
  • Marreiros J, Calandra I, Gneisinger W, Paixão E, Pedergnana A, Schunk L. 2020. Rethinking Use-Wear Analysis and Experimentation as Applied to the Study of Past Hominin Tool Use. Journal of Paleolithic Archaeology 3:475–502. DOI: https://doi.org/10.1007/s41982-020-00058-1.
  • Pedergnana A, Calandra I, Bob K, Gneisinger W, Paixão E, Schunk L, Hildebrandt A, Marreiros J. 2020. Evaluating the microscopic effect of brushing stone tools as a cleaning procedure. Quaternary International 569–570:263–276. DOI: https://doi.org/10.1016/j.quaint.2020.06.031.
  • Pedergnana A, Calandra I, Evans AA, Bob K, Hildebrandt A, Ollé A. 2020. Polish is quantitatively different on quartzite flakes used on different worked materials. PLOS ONE 15:e0243295. DOI: https://doi.org/10.1371/journal.pone.0243295.
  • Schultz JA, Anders U, Braune C, Brinkkötter JJ, Calandra I, Engels S, Findeisen E, Gailer JP, Hummel J, Jäger KRK, Kaiser TM, Kalthoff DC, von Koenigswald W, Kullmer O, Landwehr C, Mau M, Menz U, Ruf I, Schubert AM, Schulz-Kornas E, Schwermann AH, Schwermann LC, Skiba M, Steuer P, Südekum K-H, Winkler DE, Martin T. 2020. A new facet terminology for mammalian dentitions. In: Martin T, von Koenigswald W eds. Mammalian Teeth - Form and Function. Munich, Germany: Verlag Dr. Friedrich Pfeil, 11–24.
  • Schulz-Kornas E, Kaiser TM, Calandra I, Winkler DE. 2020. A brief history of quantitative wear analyses with an appeal for a holistic view on dental wear processes. In: Martin T, von Koenigswald W eds. Mammalian Teeth - Form and Function. Munich, Germany: Verlag Dr. Friedrich Pfeil, 44–53.


  • Calandra I, Pedergnana A, Gneisinger W, Marreiros J. 2019. Why should traceology learn from dental microwear, and vice-versa? Journal of Archaeological Science 110:105012. DOI: https://doi.org/10.1016/j.jas.2019.105012.
  • Calandra I, Schunk L, Bob K, Gneisinger W, Pedergnana A, Paixao E, Hildebrandt A, Marreiros J. 2019. The effect of numerical aperture on quantitative use-wear studies and its implication on reproducibility. Scientific Reports 9:6313. DOI: https://doi.org/10.1038/s41598-019-42713-w.
  • Calandra I, Schunk L, Rodriguez A, Gneisinger W, Pedergnana A, Paixao E, Pereira T, Iovita R, Marreiros J. 2019. Back to the edge: relative coordinate system for use-wear analysis. Archaeological and Anthropological Sciences 11:5937–5948. DOI: https://doi.org/10.1007/s12520-019-00801-y.


  • Eyssautier-Chuine S, Calandra I, Vaillant-Gaveau N, Fronteau G, Thomachot-Schneider C, Hubert J, Pleck J, Gommeaux M. 2018. A new preventive coating for building stones mixing a water repellent and an eco-friendly biocide. Progress in Organic Coatings 120:132–142. DOI: https://doi.org/10.1016/j.porgcoat.2018.03.022.


  • Calandra I, Labonne G, Schulz-Kornas E, Kaiser TM, Montuire S. 2016. Tooth wear as a means to quantify intra-specific variations in diet and chewing movements. Scientific Reports 6:34037. DOI: https://doi.org/10.1038/srep34037.
  • Calandra I, Merceron G. 2016. Dental microwear texture analysis in mammalian ecology. Mammal Review 46:215–228. DOI: https://doi.org/10.1111/mam.12063.
  • Calandra I, Zub K, Szafrańska PA, Zalewski A, Merceron G. 2016. Silicon-based plant defences, tooth wear and voles. Journal of Experimental Biology 219:501–507. DOI: https://doi.org/10.1242/jeb.134890.
  • Gailer JP, Calandra I, Schulz-Kornas E, Kaiser TM. 2016. Morphology is not Destiny: Discrepancy between Form, Function and Dietary Adaptation in Bovid Cheek Teeth. Journal of Mammalian Evolution 23:369–383. DOI: https://doi.org/10.1007/s10914-016-9325-1.
  • Schneider AR, Cancès B, Ponthieu M, Sobanska S, Benedetti MF, Pourret O, Conreux A, Calandra I, Martinet B, Morvan X, Gommeaux M, Marin B. 2016. Lead distribution in soils impacted by a secondary lead smelter: Experimental and modelling approaches. Science of The Total Environment 568:155–163. DOI: https://doi.org/10.1016/j.scitotenv.2016.06.001.


  • Calandra I, Labonne G, Mathieu O, Henttonen H, Lévêque J, Milloux M-J, Renvoisé É, Montuire S, Navarro N. 2015. Isotopic partitioning by small mammals in the subnivium. Ecology and Evolution 5:4132–4140. DOI: https://doi.org/10.1002/ece3.1653.


  • Schulz E, Calandra I, Kaiser TM. 2013. Feeding ecology and chewing mechanics in hoofed mammals: 3D tribology of enamel wear. Wear 300:169–179. DOI: https://doi.org/10.1016/j.wear.2013.01.115.
  • Winkler DE, van den Hoek Ostende LW, Schulz E, Calandra I, Gailer JP, Landwehr C, Kaiser TM. 2013a. Dietary divergence in space and time – Lessons from the dwarf-goat Myotragus balearicus (Pleisto-Holocene, Mallorca, Spain). Mammalian Biology 78:430–437. DOI: https://doi.org/10.1016/j.mambio.2013.08.003.
  • Winkler DE, Schulz E, Calandra I, Gailer JP, Landwehr C, Kaiser TM. 2013b. Indications for a dietary change in the extinct Bovid genus Myotragus (Plio-Holocene, Mallorca, Spain). Geobios 46:143–150. DOI: https://doi.org/10.1016/j.geobios.2012.10.010.


  • Calandra I, Schulz E, Pinnow M, Krohn S, Kaiser TM. 2012. Teasing apart the contributions of hard dietary items on 3D dental microtextures in primates. Journal of Human Evolution 63:85–98. DOI: https://doi.org/10.1016/j.jhevol.2012.05.001.


  • Calandra I, Göhlich UB, Merceron G. 2010. Feeding preferences of Gomphotherium subtapiroideum (Proboscidea, Mammalia) from the Miocene of Sandelzhausen (Northern Alpine Foreland Basin, southern Germany) through life and geological time: evidence from dental microwear analysis. Paläontologische Zeitschrift 84:205–215. DOI: https://doi.org/10.1007/s12542-010-0054-0.
  • Schulz E, Calandra I, Kaiser TM. 2010. Applying tribology to teeth of hoofed mammals. Scanning 32:162–182. DOI: https://doi.org/10.1002/sca.20181.
  • Vannier J, Calandra I, Gaillard C, Żylińska A. 2010. Priapulid worms: Pioneer horizontal burrowers at the Precambrian-Cambrian boundary. Geology 38:711–714. DOI: https://doi.org/10.1130/G30829.1.


  • Calandra I, Göhlich UB, Merceron G. 2008. How could sympatric megaherbivores coexist? Example of niche partitioning within a proboscidean community from the Miocene of Europe. Naturwissenschaften 95:831–838. DOI: https://doi.org/10.1007/s00114-008-0391-y

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